Wengrat, S.; Morales, E. A.; Wetzel, C. E.; Almeida, P. D.; Ector, L.; Bicudo, D. C. (2016). Taxonomy and ecology of Fragilaria billingsii sp. nov. and analysis of type material of Synedra rumpens var. fusa (Fragilariaceae, Bacillariophyta) from Brazil.
Phytotaxa. 270(3): 191.